Globidiscinae Okuyucu & Akbaş, 2026, subfam. nov.

Abstract

Subfamily Globidiscinae subfam. nov. urn:lsid:zoobank.org:act: 9BC6E72C-D473-47DD-9681-51754BC42BAE Type genus Globidiscus Okuyucu, 2021. Description Large miliolid tests composed of a spherical proloculus followed by an undivided tubular chamber, diversely coiled: initially glomospiral and finally planispiral involute, initially glomospiral then irregular planispiral involute/semi-evolute(?). Sometimes one or two sided ogival shape of the last whorls. No septa or pseudosepta. Lumen with common flosculinisation. The thick wall with perforations similar to comb teeth-shaped structure. Neosparitization rare to common. Umbilicus slightly depressed or absent. Aperture terminal, simple. Comparison Globidiscinae differ from Neodiscinae by the test shape of the last whorls which has one- or two-sided ogival-shaped last whorls, and is of a coiling type initially glomospiral and finally planispiral involute, initially glomospiral then irregular planispiral involute/semi-evolute(?) and with common flosculinisation; from Baisalininae and Agathammininae by a larger test and devoid of septa/pseudosepta; and from Multidiscinae by a test which has one- or two-sided ogival-shaped last whorls, a larger test and common flosculinisation. Remarks In the original description of the genus Kamurana it was tentatively attributed to the family Hemigordiopsidae Nikitina, 1969 instead of Milioliporidae Brönnimann & Zaninetti, 1971 by Altiner & Zaninetti (1977) due to the probable absence of septation and a homeomorph of the irregular forms of the genus Hemigordius Schubert, 1908. Besides, as indicated by the authors, the presence of perforations in the wall at the adult stage of the genus Kamurana makes its attribution to the Hemigordiopsidae problematic. In this study, after a detailed investigation of the wall structure of the genus Kamurana in the original description compared with recovered well-preserved material, it is clearly shown that the wall of Kamurana does not feature a perforation structure in the adult stage in the sense of authors. This structure is clearly comb teeth-shaped which developed on the porcelaneous wall related to the neosparitization. For this reason, the taxonomic position of the genus Kamurana should be located under the subfamily Globidiscinae subfam. nov. Included genera Kamurana (sensu Altiner & Zaninetti 1977); Globidiscus Okuyucu, 2021; Rectogordiopsis gen. nov., Davanella gen. nov. and Kaganella gen. nov.